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Armus and colleagues reported that Paramecium caudatum could be trained in a discrimination learning task using electrical stimulation as reinforcement.
As reported in: Armus, Montgomery, Jellison (2006). Discrimination Learning in Paramecia (P. caudatum). The Psychological Record. doi:10.1007/bf03396029
Dexter, Prabakaran and Gunawardena confirmed a 1906 report that Stentor roeseli responds to repeated stimulation with a hierarchy of bending, ciliary change, contraction and detachment, with contract-vs-detach like a fair coin toss.
As reported in: Dexter, Prabakaran, Gunawardena (2019). A Complex Hierarchy of Avoidance Behaviors in a Single-Cell Eukaryote. Current Biology. doi:10.1016/j.cub.2019.10.059